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I was just playing in DCUO and we were all trash talking on the low income people who never put a dime into any of the games they play. A few of us heard other people referring to them as "mules" so we continued the moniker since they have one for us already.
"Whales" = those who sub and/or put money into games.
"Mules" = those who play F2P games and are low income.